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A field experiment also showed that snake DNA could be detected in soil where a snake had been resting up to two weeks after its removal.

The results showed that precipitation, not temperature, was the main culprit of glacier fluctuation during around 4,500, of the past 6,000 years, or 76 percent of the time.

The research team also found that the exposed female offspring showed exaggerated digging and hyperactivity in adulthood; LR mitigated these effects.

But poorer countries angrily rejected this as too low, with the group of small island nations saying they were "deeply disappointed" with an offer that showed "contempt for our vulnerable people".

Another showed men smiling to the camera as they pointed to at least a dozen others on a boat travelling rapidly over the water.