substituted

Goalkeeper Butland - who took the captain's armband after James Tavernier was substituted - was keen to stress the collective nature of the half-time debrief.

Marler's final England appearance was against the All Blacks in July, when he broke his foot and had to be substituted after 17 minutes.

England's players, including substituted man Smith, consoled Ford as they trudged away from another painful loss.

On Duran's reaction to being substituted, Tielemans added: "He is really funny. The manager does not care. He knows everyone wants to play. It is part of the game."

The 23-year-old winger was substituted early in the second half against Greece on Thursday after picking up an injury in his right leg.