forwards

"The Y chromosome has many sections that repeat, and even some that are palindromes -- the same forwards and backwards. It makes it challenging to understand which genes are encoded there," Cothran explained.

She suffered multiple injuries when she fell forwards onto the floor and she died days later in hospital.

"I was having a taste for what else could be done, another way forwards," he said.

Gatland has opted for the more traditional balance of five forwards and three backs on his bench this time after the tactic of six forwards and two backs backfired against Fiji.

Over the past five years, the golden generation of South Africa forwards have developed arguably the most formidable and feared scrum in rugby.